| 1970 | Ph.D. (summa cum laude), University of Basel, Switzerland |
| 1971-1973 | Post-doctoral education as a visiting researcher at the University of Oxford (Prof. C.D. Darlington), and at the John Innes Institute, Norwich, UK |
| 1974-1980 | Visiting Assistant, University of Vienna |
| 1980 | ,-82,-89 Visiting Lecturer, University of Kaiserslauten, Germany |
| 1978 | Habilitation (Univ. Doz.) in Cytology, University of Vienna |
| 1983 | Appointment as Associate Professor - ao. Prof., Vienna University |
| 1986-2004 | Full Professor - o. Prof. (Cytology & Genetics in Botany) and Head of Department, Institute of Botany, University of Vienna |
| 2004-2006 | Head of Department (Chromosome Biology), Max F. Perutz Laboratories, University of Vienna |
