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Ph.D. Scott F. GILBERT Howard A. Schneiderman Professor of Biology, Department of Biology, Swarthmore College; Finland Distinguished Professor, University of Helsinki

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1971 BA, Wesleyan University, Connecticut 1976 MA, History of Biology, The Johns Hopkins University 1976 Ph.D., Biology, The Johns Hopkins University, Baltimore, Maryland 1976-1980 Postdoctoral Fellow, University of Wisconsin since 1980 Department of Biology, Swarthmore College 1983-1984 Associate Scientist, Wistar Institute of Anatomy and Biology 1987-1988 Associate Scientist, Thomas Jefferson University 1990-1991 Visiting Professor, University of Helsinki since 2009 Finland Distinguished Professorship, University of Helsinki

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